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- Irreversible Processes in Ecological Evolution/Are changes in species interactions and their ecosystem consequences irreversible? + (All interactions between individuals of th … All interactions between individuals of the same or different species (populations) are metabolically-constrained. That is, the rate of an individual's energy use (metabolic rate) sets the rate of interactions with other individuals. In this talk, I will first describe the relationship between metabolic and species interaction rates as a function of the physical environment as well as the organism's mass, using ecological metabolic theory. I will then describe the effects of (metabolically-constrained) species interactions on the dynamics of ecosystems. Finally, I will consider whether changes in metabolically-constrained species interactions are irreversible. species interactions are irreversible. )